Great apes held in captivity need resilience: the ability to ‘bounce back’ following setbacks. This is particularly true for apes entering sanctuaries or rehabilitation centres, as they have usually endured traumatic conditions, including killing of their mother and separation from their social group, which can have lasting psychological effects, impairing their resilience. This in turn threatens to reduce the effectiveness of conservation programs involving rehabilitation and release of ex-captive individuals, as reduced resilience may make it less likely that apes will be able to thrive in the wild.
However, even zoo-housed apes may need to build their resilience. One of the factors contributing to low resilience is experiencing a lack of agency. This affects captive apes in all settings because they often have no choice about when to feed and forage, what to eat, or which individuals to socialise with. Routine management practices which are important for the apes’ health or their conservation may impose stressful or frustrating situations which they cannot escape or avoid (e.g., regular veterinary health checks or treatment involving capture and sedation/anaesthesia, temporary or longer-term separation from their social group during treatment and recovery). Collection management (such as species breeding programmes) may mean that individuals are moved between institutions and must integrate into a new social group. Unlike dispersal in the wild, captive apes have no control over when this occurs and which group they join.
We will build on pilot research in our group on orangutans and gorillas housed in zoos, which has identified promising avenues for progress in addressing this problem. We have integrated research on fostering resilience in humans and other non-human animal species and used it to design interventions to build their resilience. In this project we aim to develop and test these techniques on great apes, as well as investigating how management techniques can be altered to provide apes with a sense of agency. As a result we hope to develop techniques which will build resilience, improving the welfare and conservation outcomes for great apes in zoos, sanctuaries and rehabilitation centres.
Figure 1: A captive Western lowland gorilla (Gorilla gorilla gorilla) using a stick to extract peanut butter from a puzzle feeder which has bolts blocking access. Photo: Lelia Bridgeland-Stephens.
This project is not suitable for CASE funding
Each host has a slightly different application process.
Find out how to apply for this studentship.
All applications must include the CENTA application form.
Choose your application route
DRs will be awarded CENTA Training Credits (CTCs) for participation in CENTA-provided and ‘free choice’ external training. One CTC can be earned per 3 hours training, and DRs must accrue 100 CTCs across the three and a half years of their PhD.
This PhD studentship is part of a broad research programme in which we are working with a number of partners in conservation and rehabilitation of a variety of species across the world. This includes zoos in the UK (e.g. Chester, Twycross, Drayton Manor Zoos), as well as NGOs in ape conservation in range countries (e.g. SOCP, and BOSF). We also work with the Orangutan Veterinary Advisory Group, who are a representative body for orangutan sanctuaries. Thus, the student will be embedded in a dynamic network of organisations.
Training in the specialist methodologies required for behavioural fieldwork will be provided by both supervisors. Chappell will provide training in sampling and recording behavioural indicators of welfare status, cognitive and social behaviours and in designing the experimental interventions. Thorpe will provide training recording movement and understanding ecology. Chappell and external training will provide additional training in the specific programming and statistical techniques required (e.g. R, linear modelling techniques, Bayesian analysis etc.). Both supervisors will support skills development in delivering impact and engagement from the project with zoos, sanctuaries and the public.
Not applicable.
Year 1: Literature review of the behavioural ecology of the species to be studied. Development of observational behavioural data collection on resilience indicators, and development of experimental interventions. Training in developing data collection protocols using Zoo Monitor. Initial training on data analysis and statistics. Field work at zoo(s) – initial data collection at zoos.
Year 2: Data analysis on baseline data and experimental interventions. Iteratively improve experimental interventions, depending on outcome of initial experiments, and analyse.
Year 3: After modifications to enclosures and/or management procedures, collection of post-modification dataset on behavioural indicators of resilience, and comparison to baseline data to determine if outcomes have improved. Write and publish resulting papers.
Journal:
Bridgeland-Stephens, L., Thorpe, S.K., Chappell, J., 2023. Potential resilience treatments for orangutans (Pongo spp.): Lessons from a scoping review of interventions in humans and other animals. Animal Welfare 32, e77. https://doi.org/10.1017/awf.2023.97
Thorpe, S., Neufuss, J., Myatt, J., Tarrega, E., Wamba, G., Sulistyo, F., Benítez López, A., Chappell, J., 2024. The EDT: An evidence-based framework for improving captive great ape well-being., in: Unwin, S., White, A., Landjouw, A. (Eds.), State of the Apes Vol V: Health and Disease at the Human-Ape Interface. ARCUS Foundation.
Chappell, J., Thorpe, S.K.S., 2022. The role of great ape behavioral ecology in One Health: implications for captive welfare and re-habilitation success. American Journal of Primatology 84, e23328. https://doi.org/10.1002/ajp.23328
Lyons, D.M., Parker, K.J., Schatzberg, A.F., 2010. Animal models of early life stress: implications for understanding resilience. Dev Psychobiol 52, 616–24. https://doi.org/10.1002/dev.20500
Scheffer, M., Bolhuis, J.E., Borsboom, D., Buchman, T.G., Gijzel, S.M.W., Goulson, D., Kammenga, J.E., Kemp, B., Melis, R.J.F., van Nes, E.H., Romero, L.M., 2018. Quantifying resilience of humans and other animals. Proc Natl Acad Sci USA 115, 11883–11890. https://doi.org/10.1073/pnas.1810630115
Wu, G., Feder, A., Cohen, H., Kim, J.J., Calderon, S., Charney, D.S., Mathé, A.A., 2013. Understanding resilience. Front Behav Neurosci 7, 10. https://doi.org/10.3389/fnbeh.2013.00010
We expect candidates to have a Merit or Distinction at MSc level in a relevant subject. Some experience (through a formal qualification or self-taught) of programming is essential. Experience of collecting behavioural data would also be an advantage, and field work experience would be desirable. You can find information about our project on great apes here, and a talk on our previous work on parrots here.
For any enquiries related to this project please contact
To apply to this project:
Applications must be submitted by 23:59 GMT on Wednesday 7th January 2026.